ΠΠ»ΠΈΡΠ½ΠΈΠ΅ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΡΡ Π·Π°ΠΌΠ΅ΡΠ΅Π½ΠΈΠΉ Π²Π±Π»ΠΈΠ·ΠΈ ΠΌΠΎΠ»Π΅ΠΊΡΠ» Π±Π°ΠΊΡΠ΅ΡΠΈΠΎΡ Π»ΠΎΡΠΎΡΠΈΠ»Π»ΠΎΠ² PA ΠΈ Bb Π½Π° ΡΠ²ΠΎΠΉΡΡΠ²Π° ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΠΎΠ³ΠΎ ΡΠ΅Π½ΡΡΠ° Rhodobacter sphueroides
![ΠΠΈΡΡΠ΅ΡΡΠ°ΡΠΈΡ: ΠΠ»ΠΈΡΠ½ΠΈΠ΅ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΡΡ
Π·Π°ΠΌΠ΅ΡΠ΅Π½ΠΈΠΉ Π²Π±Π»ΠΈΠ·ΠΈ ΠΌΠΎΠ»Π΅ΠΊΡΠ» Π±Π°ΠΊΡΠ΅ΡΠΈΠΎΡ
Π»ΠΎΡΠΎΡΠΈΠ»Π»ΠΎΠ² PA ΠΈ Bb Π½Π° ΡΠ²ΠΎΠΉΡΡΠ²Π° ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΠΎΠ³ΠΎ ΡΠ΅Π½ΡΡΠ° Rhodobacter sphueroides](https://niscu.ru/work/3529836/cover.png)
Π’Π΅ΠΌΠ° Π΄ΠΈΡΡΠ΅ΡΡΠ°ΡΠΈΠΈ ΡΠ²Π»ΡΠ΅ΡΡΡ ΡΠ°ΡΡΡΡ ΠΏΠ»Π°Π½ΠΎΠ²ΠΎΠΉ ΡΠ΅ΠΌΡ ΠΈΠ½ΡΡΠΈΡΡΡΠ°. Π Π°Π±ΠΎΡΠ° Π²ΡΠΏΠΎΠ»Π½ΡΠ»Π°ΡΡ ΠΏΡΠΈ ΡΠΈΠ½Π°Π½ΡΠΎΠ²ΠΎΠΉ ΠΏΠΎΠ΄Π΄Π΅ΡΠΆΠΊΠ΅ Π€Π΅Π΄Π΅ΡΠ°Π»ΡΠ½ΡΠΌ Π°Π³Π΅Π½ΡΡΡΠ²ΠΎΠΌ ΠΏΠΎ Π½Π°ΡΠΊΠ΅ ΠΈ ΠΈΠ½Π½ΠΎΠ²Π°ΡΠΈΡΠΌ Π ΠΎΡΡΠΈΠΉΡΠΊΠΎΠΉ Π°ΠΊΠ°Π΄Π΅ΠΌΠΈΠΈ Π½Π°ΡΠΊ (02.512.11.2085; 02.740.11.0293), ΠΏΡΠΎΠ³ΡΠ°ΠΌΠΌΠ°ΠΌΠΈ ΠΡΠ΅Π·ΠΈΠ΄ΠΈΡΠΌΠ° Π ΠΠ «ΠΠΎΠ»Π΅ΠΊΡΠ»ΡΡΠ½Π°Ρ ΠΈ ΠΊΠ»Π΅ΡΠΎΡΠ½Π°Ρ Π±ΠΈΠΎΠ»ΠΎΠ³ΠΈΡ», «ΠΠΊΡΡΡΠ΅ΠΌΠ°Π»ΡΠ½ΡΠ΅ ΡΠ²Π΅ΡΠΎΠ²ΡΠ΅ ΠΏΠΎΠ»Ρ ΠΈ ΠΈΡ ΠΏΡΠΈΠ»ΠΎΠΆΠ΅Π½ΠΈΠ΅», Π³ΡΠ°Π½ΡΠΎΠΌ ΠΡΠ΅Π·ΠΈΠ΄Π΅Π½ΡΠ° Π Π€ ΠΠ¨-3277.2006.4- ΠΠ¨-4525.2008.4) ΠΈ Π Π€Π€Π (03β04β48 215… Π§ΠΈΡΠ°ΡΡ Π΅ΡΡ >
- Π‘ΠΎΠ΄Π΅ΡΠΆΠ°Π½ΠΈΠ΅
- ΠΡΠ΄Π΅ΡΠΆΠΊΠ°
- ΠΠΈΡΠ΅ΡΠ°ΡΡΡΠ°
- ΠΡΡΠ³ΠΈΠ΅ ΡΠ°Π±ΠΎΡΡ
- ΠΠΎΠΌΠΎΡΡ Π² Π½Π°ΠΏΠΈΡΠ°Π½ΠΈΠΈ
Π‘ΠΎΠ΄Π΅ΡΠΆΠ°Π½ΠΈΠ΅
- Π‘ΠΏΠΈΡΠΎΠΊ ΡΡΠ»ΠΎΠ²Π½ΡΡ ΡΠΎΠΊΡΠ°ΡΠ΅Π½ΠΈΠΉ
- ΠΠΠΠΠ 1. ΠΠΠΠΠ ΠΠΠ’ΠΠ ΠΠ’Π£Π Π«
- 1. 1. ΠΠΎΠΌΠΏΠΎΠ½Π΅Π½ΡΡ ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΎΠΉ ΠΌΠ΅ΠΌΠ±ΡΠ°Π½Ρ
- 1. 2. Π€ΠΎΡΠΎΡΠΈΠ½ΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΈΠ΅ ΠΏΠΈΠ³ΠΌΠ΅Π½ΡΡ
- 1. 2. 1. ΠΠ°ΠΊΡΠ΅ΡΠΈΠΎΡ Π»ΠΎΡΠΎΡΠΈΠ»Π»Ρ ΠΈ Ρ Π»ΠΎΡΠΎΡΠΈΠ»Π»Ρ
- 1. 2. 2. Π€ΠΈΠΊΠΎΠ±ΠΈΠ»ΠΈΠ½Ρ
- 1. 2. 3. ΠΠ°ΡΠΎΡΠΈΠ½ΠΎΠΈΠ΄Ρ
- 1. 3. ΠΡΠΎΠ±Π΅Π½Π½ΠΎΡΡΠΈ ΠΊΠΎΠΎΡΠ΄ΠΈΠ½ΠΈΡΠΎΠ²Π°Π½ΠΈΡ Π°ΡΠΎΠΌΠ° ΠΌΠ°Π³Π½ΠΈΡ (Π±Π°ΠΊΡΠ΅ΡΠΈΠΎ)Ρ Π»ΠΎΡΠΎΡΠΈΠ»Π»ΠΎΠ² Π² ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΈΡ ΠΊΠΎΠΌΠΏΠ»Π΅ΠΊΡΠ°Ρ
- 1. 4. Π’ΠΈΠΏΡ, ΡΠΎΡΡΠ°Π² ΠΈ ΡΡΡΡΠΊΡΡΡΠ° ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΈΡ ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΡΡ ΡΠ΅Π½ΡΡΠΎΠ²
- 1. 5. ΠΠ΅ΡΠ΅Π½ΠΎΡ ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π° Π² Π±Π°ΠΊΡΠ΅ΡΠΈΠ°Π»ΡΠ½ΠΎΠΌ ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΠΎΠΌ ΡΠ΅Π½ΡΡΠ΅
- 1. 6. ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅ ΠΌΠ΅Ρ
Π°Π½ΠΈΠ·ΠΌΠΎΠ² ΠΏΠ΅ΡΠ΅Π½ΠΎΡΠ° ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π° Π² Π±Π°ΠΊΡΠ΅ΡΠΈΠ°Π»ΡΠ½ΡΡ
ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΡΡ
ΡΠ΅Π½ΡΡΠ°Ρ
- 1. 6. 1. ΠΠ°ΠΌΠ΅ΡΠ΅Π½ΠΈΠ΅ ΠΊΠΎΡΠ°ΠΊΡΠΎΡΠΎΠ² ΠΏΠ΅ΡΠ΅Π½ΠΎΡΠ° ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π°
- 1. 6. 2. Π‘Π°ΠΉΡ — Π½Π°ΠΏΡΠ°Π²Π»Π΅Π½Π½ΡΠΉ ΠΌΡΡΠ°Π³Π΅Π½Π΅Π· Π Π¦ ΠΏΡΡΠΏΡΡΠ½ΡΡ
Π±Π°ΠΊΡΠ΅ΡΠΈΠΉ
- 1. 6. 2. 1. ΠΡΠΏΠΎΠ»ΡΠ·ΠΎΠ²Π°Π½ΠΈΠ΅ ΠΌΠ΅ΡΠΎΠ΄ΠΎΠ² ΠΌΠΎΠ»Π΅ΠΊΡΠ»ΡΡΠ½ΠΎΠΉ Π±ΠΈΠΎΠ»ΠΎΠ³ΠΈΠΈ Π΄Π»Ρ ΠΏΠΎΠ»ΡΡΠ΅Π½ΠΈΡ Π Π¦ Rba. sphaeroides, Π² ΠΊΠΎΡΠΎΡΡΡ ΠΎΡΡΡΡΡΡΠ²ΡΡΡ ΠΊΠΎΡΠ°ΠΊΡΠΎΡΡ ΠΏΠ΅ΡΠ΅Π½ΠΎΡΠ° ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π°
- 1. 6. 2. 2. ΠΠ°ΠΌΠ΅ΡΠ΅Π½ΠΈΠ΅ ΠΊΠΎΡΠ°ΠΊΡΠΎΡΠΎΠ² ΠΏΠ΅ΡΠ΅Π½ΠΎΡΠ° ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π° Π² Π Π¦ Rba. sphaeroides ΠΏΡΠΈ ΠΏΠΎΠΌΠΎΡΠΈ ΠΌΠ΅ΡΠΎΠ΄ΠΎΠ² Π³Π΅Π½Π½ΠΎΠΉ ΠΈΠ½ΠΆΠ΅Π½Π΅ΡΠΈΠΈ
- 1. 6. 2. 3. ΠΡΡΠ°ΡΠΈΠΈ, ΠΊΠΎΡΠΎΡΡΠ΅ ΠΏΡΠΈΠ²ΠΎΠ΄ΡΡ ΠΊ ΠΈΠ·ΠΌΠ΅Π½Π΅Π½ΠΈΡ Π²ΠΎΠ΄ΠΎΡΠΎΠ΄Π½ΡΡ ΡΠ²ΡΠ·Π΅ΠΉ Π²Π±Π»ΠΈΠ·ΠΈ ΠΊΠΎΡΠ°ΠΊΡΠΎΡΠΎΠ²
- 1. 6. 2. 4. ΠΡΡΠ°ΡΠΈΠΈ, ΡΠ²ΡΠ·Π°Π½Π½ΡΠ΅ Ρ ΠΈΠ·ΠΌΠ΅Π½Π΅Π½ΠΈΠ΅ΠΌ Π°ΡΠΎΠΌΠ°ΡΠΈΡΠ΅ΡΠΊΠΈΡ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΡΡ ΠΎΡΡΠ°ΡΠΊΠΎΠ²
- 1. 6. 2. 5. ΠΠ·ΡΡΠ΅Π½ΠΈΠ΅ ΡΠΎΠ»ΠΈ ΠΊΠΎΠ½ΡΠ΅ΡΠ²Π°ΡΠΈΠ²Π½ΡΡ ΠΌΠΎΠ»Π΅ΠΊΡΠ» Π²ΠΎΠ΄Ρ Π² Π±Π°ΠΊΡΠ΅ΡΠΈΠ°Π»ΡΠ½ΠΎΠΌ Π Π¦
- 1. 6. 2. 6. ΠΡΡΠ°ΡΠΈΠΈ, ΡΠΏΠΎΡΠΎΠ±ΡΡΠ²ΡΡΡΠΈΠ΅ ΠΏΠ΅ΡΠ΅Π½ΠΎΡΡ ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π° ΠΏΠΎ Π-ΡΠ΅ΠΏΠΈ
- 1. 6. 2. 7. ΠΡΡΠ°ΡΠΈΠΈ, ΠΏΡΠΈΠ²ΠΎΠ΄ΡΡΠΈΠ΅ ΠΊ ΠΈΠ·ΠΌΠ΅Π½Π΅Π½ΠΈΡ ΡΠΈΠΌΠΌΠ΅ΡΡΠΈΠΈ Π Π¦
- 1. 6. 2. 8. Π‘ΠΎΠ·Π΄Π°Π½ΠΈΠ΅ Ρ ΠΈΠΌΠ΅ΡΠ½ΡΡ Π Π¦
- 2. 1. ΠΠ°ΡΠ΅ΡΠΈΠ°Π»Ρ
- 2. 1. 1. ΠΡΠΏΠΎΠ»ΡΠ·ΡΠ΅ΠΌΡΠ΅ Π² ΡΠ°Π±ΠΎΡΠ΅ Π±Π°ΠΊΡΠ΅ΡΠΈΠ°Π»ΡΠ½ΡΠ΅ ΡΡΠ°ΠΌΠΌΡ ΠΈ ΠΏΠ»Π°Π·ΠΌΠΈΠ΄Ρ
- 2. 1. 2. ΠΠ½ΡΠΈΠ±ΠΈΠΎΡΠΈΠΊΠΈ, ΠΈΡΠΏΠΎΠ»ΡΠ·ΡΠ΅ΠΌΡΠ΅ Π² ΡΠ°Π±ΠΎΡΠ΅
- 2. 1. 3. ΠΡΡΠ΅ΡΡ ΠΈ ΡΡΠ΅Π΄Ρ, ΠΈΡΠΏΠΎΠ»ΡΠ·ΡΠ΅ΠΌΡΠ΅ Π² ΡΠ°Π±ΠΎΡΠ΅
- 2. 2. ΠΠΈΠΊΡΠΎΠ±ΠΈΠΎΠ»ΠΎΠ³ΠΈΡΠ΅ΡΠΊΠΈΠ΅ ΠΈ ΠΌΠΎΠ»Π΅ΠΊΡΠ»ΡΡΠ½ΠΎ-Π±ΠΈΠΎΠ»ΠΎΠ³ΠΈΡΠ΅ΡΠΊΠΈΠ΅ ΠΌΠ΅ΡΠΎΠ΄Ρ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ
- 2. 2. 1. ΠΡΡΠ°ΡΠΈΠ²Π°Π½ΠΈΠ΅ Π±Π°ΠΊΡΠ΅ΡΠΈΠ°Π»ΡΠ½ΡΡ ΡΡΠ°ΠΌΠΌΠΎΠ²
- 2. 2. 2. ΠΡΠ΄Π΅Π»Π΅Π½ΠΈΠ΅ ΡΠΎΡΠ°Π»ΡΠ½ΠΎΠΉ ΠΠΠ Rba.sphaeroid.es
- 2. 2. 3. ΠΡΠ΄Π΅Π»Π΅Π½ΠΈΠ΅ ΠΏΠ»Π°Π·ΠΌΠΈΠ΄Π½ΠΎΠΉ ΠΠΠ
- 2. 2. 4. ΠΠΎΠ»ΠΈΠΌΠ΅ΡΠ°Π·Π½Π°Ρ ΡΠ΅ΠΏΠ½Π°Ρ ΡΠ΅Π°ΠΊΡΠΈΡ
- 2. 2. 5. ΠΠΎΡΠΈΠ·ΠΎΠ½ΡΠ°Π»ΡΠ½ΡΠΉ ΡΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅Π· Π² Π°Π³Π°ΡΠΎΠ·Π½ΠΎΠΌ Π³Π΅Π»Π΅
- 2. 2. 6. ΠΡΠ΅ΠΏΠ°ΡΠ°ΡΠΈΠ²Π½ΠΎΠ΅ Π²ΡΠ΄Π΅Π»Π΅Π½ΠΈΠ΅ ΡΡΠ°Π³ΠΌΠ΅Π½ΡΠΎΠ² ΠΠΠ ΠΈΠ· Π»Π΅Π³ΠΊΠΎΠΏΠ»Π°Π²ΠΊΠΎΠΉ Π°Π³Π°ΡΠΎΠ·Ρ
- 2. 2. 7. ΠΠΈΠ³ΠΈΡΠΎΠ²Π°Π½ΠΈΠ΅ ΠΌΠΎΠ»Π΅ΠΊΡΠ»Ρ Π²Π΅ΠΊΡΠΎΡΠ° ΠΈ ΡΡΠ°Π³ΠΌΠ΅Π½ΡΠ°
- 2. 2. 8. ΠΠ»Π΅ΠΊΡΡΠΎΠΏΠΎΡΠ°ΡΠΈΡ
- 2. 2. 9. ΠΠ΅ΡΠ΅Π½ΠΎΡ ΠΏΠ»Π°Π·ΠΌΠΈΠ΄ Π΄Π»Ρ ΠΊΠΎΠΌΠΏΠ»Π΅ΠΌΠ΅Π½ΡΠ°ΡΠΈΠΈ ΠΈΠ· Π. coli Π² Rba. sphaeroides
- 2. 3. ΠΠ΅ΡΠΎΠ΄Ρ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΡΡ
ΡΠ΅Π½ΡΡΠΎΠ²
- 2. 3. 1. ΠΡΠ΄Π΅Π»Π΅Π½ΠΈΠ΅ ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΡΡ ΡΠ΅Π½ΡΡΠΎΠ²
- 2. 3. 2. ΠΠ·ΠΌΠ΅ΡΠ΅Π½ΠΈΠ΅ ΡΠΏΠ΅ΠΊΡΡΠΎΠ² ΠΏΠΎΠ³Π»ΠΎΡΠ΅Π½ΠΈΡ Π Π¦
- 2. 3. 3. ΠΠ·ΠΌΠ΅ΡΠ΅Π½ΠΈΠ΅ Π΄ΠΈΡΡΠ΅ΡΠ΅Π½ΡΠΈΠ°Π»ΡΠ½ΡΡ (ΡΠ²Π΅Ρ-ΠΌΠΈΠ½ΡΡ-ΡΠ΅ΠΌΠ½ΠΎΡΠ°) ΡΠΏΠ΅ΠΊΡΡΠΎΠ²
- 2. 3. 4. ΠΠΏΡΠ΅Π΄Π΅Π»Π΅Π½ΠΈΠ΅ ΠΏΠΈΠ³ΠΌΠ΅Π½ΡΠ½ΠΎΠ³ΠΎ ΡΠΎΡΡΠ°Π²Π°
- 2. 3. 5. ΠΠ·ΠΌΠ΅ΡΠ΅Π½ΠΈΠ΅ Π΄ΠΈΡΡΠ΅ΡΠ΅Π½ΡΠΈΠ°Π»ΡΠ½ΡΡ (ΡΠ²Π΅Ρ-ΠΌΠΈΠ½ΡΡ-ΡΠ΅ΠΌΠ½ΠΎΡΠ°) ΠΠ-Π€Π’ ΡΠΏΠ΅ΠΊΡΡΠΎΠ²
- 2. 3. 6. ΠΠ·ΠΌΠ΅ΡΠ΅Π½ΠΈΠ΅ ΠΎΠΊΠΈΡΠ»ΠΈΡΠ΅Π»ΡΠ½ΠΎ-Π²ΠΎΡΡΡΠ°Π½ΠΎΠ²ΠΈΡΠ΅Π»ΡΠ½ΠΎΠ³ΠΎ ΠΏΠΎΡΠ΅Π½ΡΠΈΠ°Π»Π° ΠΏΠ΅ΡΠ²ΠΈΡΠ½ΠΎΠ³ΠΎ Π΄ΠΎΠ½ΠΎΡΠ° ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π° Π
- 2. 3. 7. ΠΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅Π· Π±Π΅Π»ΠΊΠΎΠ² Π² ΠΏΠΎΠ»ΠΈΠ°ΠΊΡΠΈΠ»Π°ΠΌΠΈΠ΄Π½ΠΎΠΌ Π³Π΅Π»Π΅ Π² Π΄Π΅Π½Π°ΡΡΡΠΈΡΡΡΡΠΈΡ ΡΡΠ»ΠΎΠ²ΠΈΡΡ
- 2. 3. 8. ΠΠ±ΡΠ°Π±ΠΎΡΠΊΠ° Π Π¦ Π±ΠΎΡΠ³ΠΈΠ΄ΡΠΈΠ΄ΠΎΠΌ Π½Π°ΡΡΠΈΡ
- 2. 3. 9. ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅ ΡΡΠ°Π±ΠΈΠ»ΡΠ½ΠΎΡΡΠΈ Π Π¦
- 2. 3. 10. ΠΠ±ΡΠ°Π±ΠΎΡΠΊΠ° Π Π¦ I (L177)H ΠΏΡΠΎΡΠ΅Π°Π·Π°ΠΌΠΈ
- 2. 3. 11. ΠΡΠ³ΠΊΠ°Ρ Π΄Π΅Π½Π°ΡΡΡΠ°ΡΠΈΡ ΠΌΡΡΠ°Π½ΡΠ½ΡΡ Π Π¦ ΠΏΡΠΈ ΠΏΠΎΠΌΠΎΡΠΈ 8 Π ΡΠ°ΡΡΠ²ΠΎΡΠ° ΠΌΠΎΡΠ΅Π²ΠΈΠ½Ρ
- 2. 3. 12. ΠΠ·ΠΌΠ΅ΡΠ΅Π½ΠΈΠ΅ ΡΠΏΠ΅ΠΊΡΡΠΎΠ² ΠΠΠ
- 2. 3. 13. Π€Π΅ΠΌΡΠΎΡΠ΅ΠΊΡΠ½Π΄Π½Π°Ρ ΡΠΏΠ΅ΠΊΡΡΠΎΡΠΊΠΎΠΏΠΈΡ Π Π¦ Π΄ΠΈΠΊΠΎΠ³ΠΎ ΡΠΈΠΏΠ° ΠΈ Π³Π΅Π½Π΅ΡΠΈΡΠ΅ΡΠΊΠΈ ΠΌΠΎΠ΄ΠΈΡΠΈΡΠΈΡΠΎΠ²Π°Π½Π½ΡΡ Π Π¦ I (L177)H ΠΈ I (L177)H+H (L173)L
- 3. 1. ΠΠ½Π΅ΡΠ΅Π½ΠΈΠ΅ Π½Π°ΠΏΡΠ°Π²Π»Π΅Π½Π½ΡΡ
Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΡΡ
Π·Π°ΠΌΠ΅ΡΠ΅Π½ΠΈΠΉ Π² Π Π¦ Rba. sphaeroides
- 3. 1. 1. ΠΠ½Π΅ΡΠ΅Π½ΠΈΠ΅ Π½Π°ΠΏΡΠ°Π²Π»Π΅Π½Π½ΠΎΠ³ΠΎ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΠΎΠ³ΠΎ Π·Π°ΠΌΠ΅ΡΠ΅Π½ΠΈΡ I (L177)H Π² Π Π¦Π―Π¬Ρ. sphaeroides
- 3. 1. 2. ΠΠ½Π΅ΡΠ΅Π½ΠΈΠ΅ Π½Π°ΠΏΡΠ°Π²Π»Π΅Π½Π½ΡΡ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΡΡ Π·Π°ΠΌΠ΅ΡΠ΅Π½ΠΈΠΉ H (L 173) L+I (L 177) Π Π² Π Π¦ Rba. sphaeroides
- 3. 1. 3. ΠΠ½Π΅ΡΠ΅Π½ΠΈΠ΅ Π½Π°ΠΏΡΠ°Π²Π»Π΅Π½Π½ΡΡ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΡΡ Π·Π°ΠΌΠ΅ΡΠ΅Π½ΠΈΠΉ I (L 177) Π+Π (Π182)L Π² Π Π¦ Rba. sphaeroides
- 3. 2. ΠΡΠΎΠ±Π΅Π½Π½ΠΎΡΡΠΈ Π²ΡΠ΄Π΅Π»Π΅Π½ΠΈΡ ΠΈ ΠΎΡΠΈΡΡΠΊΠΈ Π Π¦ Ρ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΡΠΌΠΈ Π·Π°ΠΌΠ΅Π½Π°ΠΌΠΈ I (L177)H- I (L 177) H+H (L 173) L- I (L177)H+H (M182)L
- 3. 3. Π‘ΠΏΠ΅ΠΊΡΡΡ ΠΏΠΎΠ³Π»ΠΎΡΠ΅Π½ΠΈΡ ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΡΡ ΡΠ΅Π½ΡΡΠΎΠ² I (L177)H- I (L 177) H+H (L 173) L- I (L177)H+H (M182)L, ΠΈΠ·ΠΌΠ΅ΡΠ΅Π½Π½ΡΠ΅ ΠΏΡΠΈ ΠΊΠΎΠΌΠ½Π°ΡΠ½ΠΎΠΉ ΡΠ΅ΠΌΠΏΠ΅ΡΠ°ΡΡΡΠ΅
- 3. 4. ΠΠΈΡΡΠ΅ΡΠ΅Π½ΡΠΈΠ°Π»ΡΠ½ΡΠ΅ (ΡΠ²Π΅Ρ ΠΌΠΈΠ½ΡΡ ΡΠ΅ΠΌΠ½ΠΎΡΠ°) ΡΠΏΠ΅ΠΊΡΡΡ ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΡΡ ΡΠ΅Π½ΡΡΠΎΠ² I (L177)H- I (L 177) H+H (L 173) L- I (L177)H+H (M182)L, ΠΈΠ·ΠΌΠ΅ΡΠ΅Π½Π½ΡΠ΅ ΠΏΡΠΈ ΠΊΠΎΠΌΠ½Π°ΡΠ½ΠΎΠΉ ΡΠ΅ΠΌΠΏΠ΅ΡΠ°ΡΡΡΠ΅
- 3. 5. Π‘ΠΏΠ΅ΠΊΡΡΡ ΠΏΠΎΠ³Π»ΠΎΡΠ΅Π½ΠΈΡ ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΡΡ ΡΠ΅Π½ΡΡΠΎΠ² I (L177)H- I (L 177) H+H (L 173) L- I (L177)H+H (M182)L, ΠΈΠ·ΠΌΠ΅ΡΠ΅Π½Π½ΡΠ΅ ΠΏΡΠΈ Π½ΠΈΠ·ΠΊΠΈΡ ΡΠ΅ΠΌΠΏΠ΅ΡΠ°ΡΡΡΠ°Ρ
- 3. 6. ΠΠΈΠ³ΠΌΠ΅Π½ΡΠ½ΡΠΉ Π°Π½Π°Π»ΠΈΠ· Π Π¦ Π΄ΠΈΠΊΠΎΠ³ΠΎ ΡΠΈΠΏΠ° ΠΈ Π³Π΅Π½Π΅ΡΠΈΡΠ΅ΡΠΊΠΈ ΠΌΠΎΠ΄ΠΈΡΠΈΡΠΈΡΠΎΠ²Π°Π½Π½ΡΡ Π Π¦
- 3. 7. Π‘ΠΏΠ΅ΠΊΡΡΡ ΠΏΠΎΠ³Π»ΠΎΡΠ΅Π½ΠΈΡ Π Π¦ Π΄ΠΈΠΊΠΎΠ³ΠΎ ΡΠΈΠΏΠ° ΠΈ I (L177)H, I (L177)H+H (L173)L, I (L177)H+H (M182)L ΠΏΠΎΡΠ»Π΅ ΡΠΊΡΡΡΠ°Π³ΠΈΡΠΎΠ²Π°Π½ΠΈΡ ΠΏΠΈΠ³ΠΌΠ΅Π½ΡΠΎΠ²
- 3. 8. ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅ Π Π¦ Ρ ΠΏΡΠΎΡΠ½ΠΎΡΠ²ΡΠ·Π°Π½Π½ΠΎΠΉ ΠΌΠΎΠ»Π΅ΠΊΡΠ»ΠΎΠΉ ΠΠ₯Π» ΠΌΠ΅ΡΠΎΠ΄ΠΎΠΌ ΡΠ»Π΅ΠΊΡΡΠΎΡΠΎΡΠ΅Π·Π° Π² Π΄Π΅Π½Π°ΡΡΡΠΈΡΡΡΡΠΈΡ ΡΡΠ»ΠΎΠ²ΠΈΡΡ
- 3. 9. ΠΠ·ΠΌΠ΅ΡΠ΅Π½ΠΈΠ΅ ΠΎΠΊΠΈΡΠ»ΠΈΡΠ΅Π»ΡΠ½ΠΎ-Π²ΠΎΡΡΡΠ°Π½ΠΎΠ²ΠΈΡΠ΅Π»ΡΠ½ΠΎΠ³ΠΎ ΠΏΠΎΡΠ΅Π½ΡΠΈΠ°Π»Π° Π Π¦ Π΄ΠΈΠΊΠΎΠ³ΠΎ ΡΠΈΠΏΠ° ΠΈ I (L177)H, I (L177)H+H (L173)L
- 3. 10. ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅ ΡΠ²ΠΎΠΉΡΡΠ² Π Π¦ I (L177)H
- 3. 10. 1. ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅ ΡΡΠ°Π±ΠΈΠ»ΡΠ½ΠΎΡΡΠΈ Π Π¦ I (L177)H
- 3. 10. 2. ΠΠ΅ΡΠ²ΠΈΡΠ½ΡΠ΅ ΠΏΡΠΎΡΠ΅ΡΡΡ ΡΠ°Π·Π΄Π΅Π»Π΅Π½ΠΈΡ Π·Π°ΡΡΠ΄ΠΎΠ² Π² Π Π¦ I (L177)H
- 3. 10. 3. ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅ ΡΠΎΡΡΠΎΡΠ½ΠΈΡ Π + ΠΌΠ΅ΡΠΎΠ΄ΠΎΠΌ ΠΠΠ
- 3. 11. ΠΠΏΡΠ΅Π΄Π΅Π»Π΅Π½ΠΈΠ΅ ΠΏΠΎΠ»ΠΎΠΆΠ΅Π½ΠΈΡ ΠΠ₯Π», ΠΊΠΎΠ²Π°Π»Π΅Π½ΡΠ½ΠΎ ΡΠ²ΡΠ·Π°Π½Π½ΠΎΠ³ΠΎ Ρ Π±Π΅Π»ΠΊΠΎΠΌ
- 3. 12. ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅ ΡΠ²ΠΎΠΉΡΡΠ² ΠΠ₯Π», ΠΊΠΎΠ²Π°Π»Π΅Π½ΡΠ½ΠΎ ΡΠ²ΡΠ·Π°Π½Π½ΠΎΠ³ΠΎ Ρ Π±Π΅Π»ΠΊΠΎΠ²ΠΎΠΉ ΡΡΠ±ΡΠ΅Π΄ΠΈΠ½ΠΈΡΠ΅ΠΉ
- 3. 12. 1. ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅ ΡΡΠ°Π±ΠΈΠ»ΡΠ½ΠΎΡΡΠΈ ΡΠ²ΡΠ·ΠΈ ΠΏΡΠΈ ΠΊΠΈΡΠ»ΡΡ ΠΈ Π½Π΅ΠΉΡΡΠ°Π»ΡΠ½ΡΡ Π·Π½Π°ΡΠ΅Π½ΠΈΡΡ ΡΠ
- 3. 12. 2. ΠΠ±ΡΠ°Π±ΠΎΡΠΊΠ° ΠΏΡΠΎΡΠ΅Π°Π·Π°ΠΌΠΈ Π±Π΅Π»ΠΊΠ° Π Π¦ Ρ ΠΊΠΎΠ²Π°Π»Π΅Π½ΡΠ½ΠΎ ΡΠ²ΡΠ·Π°Π½Π½ΡΠΌ ΠΠ₯Π»
- 3. 12. 3. ΠΠ·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΠΈΠ΅ ΠΊΠΎΠ²Π°Π»Π΅Π½ΡΠ½ΠΎ ΡΠ²ΡΠ·Π°Π½Π½ΠΎΠ³ΠΎ Ρ Π±Π΅Π»ΠΊΠΎΠΌ ΠΠ₯Π» Ρ Π±ΠΎΡΠ³ΠΈΠ΄ΡΠΈΠ΄ΠΎΠΌ Π½Π°ΡΡΠΈΡ
- 3. 12. 4. ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅ ΡΠΏΠ΅ΠΊΡΡΠ°Π»ΡΠ½ΡΡ ΡΠ²ΠΎΠΉΡΡΠ² ΠΠ₯Π», ΠΊΠΎΠ²Π°Π»Π΅Π½ΡΠ½ΠΎ ΡΠ²ΡΠ·Π°Π½Π½ΠΎΠ³ΠΎ Ρ Π±Π΅Π»ΠΊΠΎΠΌ Π Π¦
- 3. 12. 5. ΠΠΠ ΡΠΏΠ΅ΠΊΡΡΡ ΡΡΠΈΠΏΠ»Π΅ΡΠ½ΠΎΠ³ΠΎ ΡΠΎΡΡΠΎΡΠ½ΠΈΡ ΡΠ²ΠΎΠ±ΠΎΠ΄Π½ΠΎΠ³ΠΎ ΠΠ₯Π» ΠΈ ΠΠ₯Π», ΡΠ²ΡΠ·Π°Π½Π½ΠΎΠ³ΠΎ Ρ Π±Π΅Π»ΠΊΠΎΠΌ
- 3. 13. ΠΠ·ΠΌΠ΅ΡΠ΅Π½ΠΈΠ΅ Π΄ΠΈΡΡΠ΅ΡΠ΅Π½ΡΠΈΠ°Π»ΡΠ½ΡΡ ΠΠ-Π€Π’ ΡΠΏΠ΅ΠΊΡΡΠΎΠ² Π Π¦ Π΄ΠΈΠΊΠΎΠ³ΠΎ ΡΠΈΠΏΠ° ΠΈ I (L 177) H+H (L 173) L
- 3. 14. ΠΠ΅ΡΠ²ΠΈΡΠ½ΡΠ΅ ΠΏΡΠΎΡΠ΅ΡΡΡ ΡΠ°Π·Π΄Π΅Π»Π΅Π½ΠΈΡ Π·Π°ΡΡΠ΄ΠΎΠ² Π² Π Π¦ I (L177)H+H (L173)L.l
ΠΠ»ΠΈΡΠ½ΠΈΠ΅ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΡΡ Π·Π°ΠΌΠ΅ΡΠ΅Π½ΠΈΠΉ Π²Π±Π»ΠΈΠ·ΠΈ ΠΌΠΎΠ»Π΅ΠΊΡΠ» Π±Π°ΠΊΡΠ΅ΡΠΈΠΎΡ Π»ΠΎΡΠΎΡΠΈΠ»Π»ΠΎΠ² PA ΠΈ Bb Π½Π° ΡΠ²ΠΎΠΉΡΡΠ²Π° ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΠΎΠ³ΠΎ ΡΠ΅Π½ΡΡΠ° Rhodobacter sphueroides (ΡΠ΅ΡΠ΅ΡΠ°Ρ, ΠΊΡΡΡΠΎΠ²Π°Ρ, Π΄ΠΈΠΏΠ»ΠΎΠΌ, ΠΊΠΎΠ½ΡΡΠΎΠ»ΡΠ½Π°Ρ)
ΠΠΊΡΡΠ°Π»ΡΠ½ΠΎΡΡΡ ΡΠ΅ΠΌΡ
.
ΠΠΊΡΡΠ°Π»ΡΠ½ΠΎΡΡΡ ΠΈΠ·ΡΡΠ΅Π½ΠΈΡ ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅Π·Π° ΡΠ²ΡΠ·Π°Π½Π° Ρ ΡΠ΅ΠΌ, ΡΡΠΎ ΡΡΠΎΡ ΠΏΡΠΎΡΠ΅ΡΡ ΠΎΠ±Π΅ΡΠΏΠ΅ΡΠΈΠ²Π°Π΅Ρ Π²ΠΎΠ·ΠΌΠΎΠΆΠ½ΠΎΡΡΡ ΡΡΡΠ΅ΡΡΠ²ΠΎΠ²Π°Π½ΠΈΡ ΠΆΠΈΠ·Π½ΠΈ Π½Π° ΠΠ΅ΠΌΠ»Π΅. ΠΠ·Π²Π΅ΡΡΠ½ΠΎ, ΡΡΠΎ ΠΎΠ΄Π½ΠΈΠΌ ΠΈΠ· ΠΊΠ»ΡΡΠ΅Π²ΡΡ ΡΡΠ°ΠΏΠΎΠ² ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅Π·Π° ΡΠ²Π»ΡΡΡΡΡ ΠΏΠ΅ΡΠ²ΠΈΡΠ½ΡΠ΅ ΡΠΎΡΠΎΠΏΡΠΎΡΠ΅ΡΡΡ ΡΠ΅Π°ΠΊΡΠΈΠΈ, Π² ΠΊΠΎΡΠΎΡΡΡ ΡΠ½Π΅ΡΠ³ΠΈΡ ΡΠ²Π΅ΡΠ°, ΠΏΠΎΠ³Π»ΠΎΡΠ΅Π½Π½Π°Ρ ΠΏΠΈΠ³ΠΌΠ΅Π½ΡΠ°ΠΌΠΈ ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅Π·ΠΈΡΡΡΡΠ΅Π³ΠΎ ΠΎΡΠ³Π°Π½ΠΈΠ·ΠΌΠ°, ΠΏΡΠ΅ΠΎΠ±ΡΠ°Π·ΡΠ΅ΡΡΡ Π² ΡΠ½Π΅ΡΠ³ΠΈΡ Ρ ΠΈΠΌΠΈΡΠ΅ΡΠΊΠΈΡ ΡΠ²ΡΠ·Π΅ΠΉ. ΠΠ°ΡΠ°Π»ΡΠ½ΡΠ΅ ΡΡΠ°ΠΏΡ ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅Π·Π° ΠΌΠΎΠΆΠ½ΠΎ Π½Π°Π·Π²Π°ΡΡ «ΡΠ½Π΅ΡΠ³Π΅ΡΠΈΡΠ΅ΡΠΊΠΈΠΌΠΈ Π²ΠΎΡΠΎΡΠ°ΠΌΠΈ ΠΆΠΈΠ·Π½ΠΈ», ΡΠ°ΠΊ ΠΊΠ°ΠΊ ΠΈΠΌΠ΅Π½Π½ΠΎ ΡΠ°ΠΌ ΠΏΡΠΎΠΈΡΡ ΠΎΠ΄ΠΈΡ Π½Π΅ΠΏΠΎΡΡΠ΅Π΄ΡΡΠ²Π΅Π½Π½ΠΎΠ΅ Π²Π·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΠΈΠ΅ ΡΠΈΠ·ΠΈΡΠ΅ΡΠΊΠΈΡ ΡΠ²Π»Π΅Π½ΠΈΠΉ (ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π½ΠΎΠ³ΠΎ Π²ΠΎΠ·Π±ΡΠΆΠ΄Π΅Π½ΠΈΡ, ΠΏΠ΅ΡΠ΅Π½ΠΎΡΠ° ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π°) Ρ Π±ΠΈΠΎΠ»ΠΎΠ³ΠΈΡΠ΅ΡΠΊΠΈΠΌΠΈ. ΠΡΠ΅ΠΎΠ±ΡΠ°Π·ΠΎΠ²Π°Π½ΠΈΠ΅ ΡΠ½Π΅ΡΠ³ΠΈΠΈ ΡΠ²Π΅ΡΠ° ΠΏΡΠΎΠΈΡΡ ΠΎΠ΄ΠΈΡ Π² ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΈΡ ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΡΡ ΡΠ΅Π½ΡΡΠ°Ρ (Π Π¦). Π Π¦ ΠΏΡΡΠΏΡΡΠ½ΠΎΠΉ Π±Π°ΠΊΡΠ΅ΡΠΈΠΈ Rhodobacter (Rba.) sphaeroides ΠΏΡΠ΅Π΄ΡΡΠ°Π²Π»ΡΠ΅Ρ ΡΠΎΠ±ΠΎΠΉ ΡΡΠ°Π½ΡΠΌΠ΅ΠΌΠ±ΡΠ°Π½Π½ΡΠΉ ΠΏΠΈΠ³ΠΌΠ΅Π½Ρ-Π±Π΅Π»ΠΊΠΎΠ²ΡΠΉ ΠΊΠΎΠΌΠΏΠ»Π΅ΠΊΡ, ΡΠΎΡΡΠΎΡΡΠΈΠΉ ΠΈΠ· ΡΡΠ΅Ρ Π±Π΅Π»ΠΊΠΎΠ²ΡΡ ΡΡΠ±ΡΠ΅Π΄ΠΈΠ½ΠΈΡ (Π, Π ΠΈ L) ΠΈ Π΄Π΅ΡΡΡΠΈ ΠΊΠΎΡΠ°ΠΊΡΠΎΡΠΎΠ² ΠΏΠ΅ΡΠ΅Π½ΠΎΡΠ° ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π°. Π‘ΡΡΡΠΊΡΡΡΠ° ΡΡΠΎΠ³ΠΎ Π Π¦ Π²ΠΏΠ΅ΡΠ²ΡΠ΅ Π±ΡΠ»Π° ΠΎΠΏΡΠ΅Π΄Π΅Π»Π΅Π½Π° Π² 1984 Π³. Π² Π»Π°Π±ΠΎΡΠ°ΡΠΎΡΠΈΠΈ Michel, ΠΠ΅ΡΠΌΠ°Π½ΠΈΡ, ΠΎΡΠΊΡΡΠ² Π²ΠΎΠ·ΠΌΠΎΠΆΠ½ΠΎΡΡΠΈ Π΄Π»Ρ Π΄Π΅ΡΠ°Π»ΡΠ½ΠΎΠ³ΠΎ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ ΠΌΠ΅Ρ Π°Π½ΠΈΠ·ΠΌΠΎΠ² ΠΏΠ΅ΡΠ²ΠΈΡΠ½ΡΡ ΡΡΠ°Π΄ΠΈΠΉ ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅Π·Π°. Π€ΠΎΡΠΎΠΈΠ½Π΄ΡΡΠΈΡΠΎΠ²Π°Π½Π½ΠΎΠ΅ ΡΠ°Π·Π΄Π΅Π»Π΅Π½ΠΈΠ΅ Π·Π°ΡΡΠ΄ΠΎΠ² Π² Π Π¦ ΡΠ²Π»ΡΠ΅ΡΡΡ ΡΠ²Π΅ΡΡ Π±ΡΡΡΡΡΠΌ ΠΏΡΠΎΡΠ΅ΡΡΠΎΠΌ, ΠΊΠ²Π°Π½ΡΠΎΠ²ΡΠΉ Π²ΡΡ ΠΎΠ΄ ΠΊΠΎΡΠΎΡΠΎΠ³ΠΎ Π±Π»ΠΈΠ·ΠΎΠΊ ΠΊ Π΅Π΄ΠΈΠ½ΠΈΡΠ΅. Π‘ΠΎΠ³Π»Π°ΡΠ½ΠΎ ΡΠΎΠ²ΡΠ΅ΠΌΠ΅Π½Π½ΡΠΌ ΠΏΡΠ΅Π΄ΡΡΠ°Π²Π»Π΅Π½ΠΈΡΠΌ, Π²ΡΡΠΎΠΊΠ°Ρ ΡΡΡΠ΅ΠΊΡΠΈΠ²Π½ΠΎΡΡΡ ΡΡΠ½ΠΊΡΠΈΠΎΠ½ΠΈΡΠΎΠ²Π°Π½ΠΈΡ Π Π¦ ΠΎΠΏΡΠ΅Π΄Π΅Π»ΡΠ΅ΡΡΡ ΡΠ²ΠΎΠΉΡΡΠ²Π°ΠΌΠΈ ΠΊΠΎΡΠ°ΠΊΡΠΎΡΠΎΠ², ΠΈΡ Π²Π·Π°ΠΈΠΌΠΎΠ΄Π΅ΠΉΡΡΠ²ΠΈΠ΅ΠΌ Π΄ΡΡΠ³ Ρ Π΄ΡΡΠ³ΠΎΠΌ ΠΈ Ρ Π±Π΅Π»ΠΊΠΎΠ²ΡΠΌ ΠΎΠΊΡΡΠΆΠ΅Π½ΠΈΠ΅ΠΌ. ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΡΡ ΡΠ΅Π½ΡΡΠΎΠ² ΠΏΡΡΠΏΡΡΠ½ΡΡ Π±Π°ΠΊΡΠ΅ΡΠΈΠΉ Π²Π½Π΅ΡΠ»ΠΈ ΠΎΠ³ΡΠΎΠΌΠ½ΡΠΉ Π²ΠΊΠ»Π°Π΄ Π² ΠΏΠΎΠ½ΠΈΠΌΠ°Π½ΠΈΠ΅ ΠΌΠ΅Ρ Π°Π½ΠΈΠ·ΠΌΠΎΠ² ΡΡΠ°Π½ΡΡΠΎΡΠΌΠ°ΡΠΈΠΈ ΡΠ½Π΅ΡΠ³ΠΈΠΈ ΠΏΡΠΈ ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅Π·Π΅ ΠΈ ΡΡΠ²Π΅ΡΠ΄ΠΈΠ»ΠΈ Π±Π°ΠΊΡΠ΅ΡΠΈΠ°Π»ΡΠ½ΡΠ΅ Π Π¦ Π² ΠΊΠ°ΡΠ΅ΡΡΠ²Π΅ Ρ ΠΎΡΠΎΡΠ΅ΠΉ ΠΌΠΎΠ΄Π΅Π»ΠΈ Π΄Π»Ρ ΠΈΠ·ΡΡΠ΅Π½ΠΈΡ ΠΏΠ΅ΡΠ΅Π½ΠΎΡΠ° ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π° Π² Π±ΠΈΠΎΠ»ΠΎΠ³ΠΈΡΠ΅ΡΠΊΠΈΡ ΡΠΈΡΡΠ΅ΠΌΠ°Ρ . ΠΠ΄Π½ΠΈΠΌ ΠΈΠ· ΠΏΠΎΠ΄Ρ ΠΎΠ΄ΠΎΠ² ΠΊ ΠΈΠ·ΡΡΠ΅Π½ΠΈΡ ΡΠΎΠ»ΠΈ Π±Π΅Π»ΠΊΠΎΠ²ΠΎΠ³ΠΎ ΠΎΠΊΡΡΠΆΠ΅Π½ΠΈΡ Π² ΡΡΠ½ΠΊΡΠΈΠΎΠ½ΠΈΡΠΎΠ²Π°Π½ΠΈΠΈ Π Π¦ ΡΠ²Π»ΡΠ΅ΡΡΡ Π½Π°ΠΏΡΠ°Π²Π»Π΅Π½Π½ΡΠΉ ΠΌΡΡΠ°Π³Π΅Π½Π΅Π·, Π° ΠΈΠΌΠ΅Π½Π½ΠΎ ΡΠ΅Π»Π΅ΠΊΡΠΈΠ²Π½ΠΎΠ΅ Π·Π°ΠΌΠ΅ΡΠ΅Π½ΠΈΠ΅ ΠΊΠΎΠ½ΡΠ΅ΡΠ²Π°ΡΠΈΠ²Π½ΡΡ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΡΡ ΠΎΡΡΠ°ΡΠΊΠΎΠ² Π² ΡΡΠ°ΡΡΠΊΠ°Ρ ΡΠ²ΡΠ·ΡΠ²Π°Π½ΠΈΡ Ρ ΡΠΎΠΌΠΎΡΠΎΡΠΎΠ². Π‘ΠΎΡΠ΅ΡΠ°Π½ΠΈΠ΅ ΡΡΠΎΠ³ΠΎ ΠΌΠ΅ΡΠΎΠ΄Π° Ρ ΡΠΎΠ²ΡΠ΅ΠΌΠ΅Π½Π½ΡΠΌΠΈ Π±ΠΈΠΎΡ ΠΈΠΌΠΈΡΠ΅ΡΠΊΠΈΠΌΠΈ ΠΈ Π±ΠΈΠΎΡΠΈΠ·ΠΈΡΠ΅ΡΠΊΠΈΠΌΠΈ ΠΌΠ΅ΡΠΎΠ΄Π°ΠΌΠΈ ΡΠ²Π»ΡΠ΅ΡΡΡ ΠΏΠ΅ΡΡΠΏΠ΅ΠΊΡΠΈΠ²Π½ΡΠΌ ΠΏΠΎΠ΄Ρ ΠΎΠ΄ΠΎΠΌ ΠΊ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ ΠΌΠ΅Ρ Π°Π½ΠΈΠ·ΠΌΠΎΠ² Π½Π°ΡΠ°Π»ΡΠ½ΡΡ ΡΡΠ°Π΄ΠΈΠΉ ΡΠΎΡΠΎΠΏΠ΅ΡΠ΅Π½ΠΎΡΠ° ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π° Π² Π Π¦ ΠΈ ΠΈΠ·ΡΡΠ΅Π½ΠΈΡ ΡΠΎΠ»ΠΈ Π±Π΅Π»ΠΊΠ° Π² ΡΡΠΎΠΌ ΠΏΡΠΎΡΠ΅ΡΡΠ΅.
Π¦Π΅Π»Ρ ΡΠ°Π±ΠΎΡΡ ΠΈ Π·Π°Π΄Π°ΡΠΈ ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ.
Π¦Π΅Π»ΡΡ ΡΠ°Π±ΠΎΡΡ Π±ΡΠ»ΠΎ Π²ΡΡΡΠ½Π΅Π½ΠΈΠ΅ Π²Π»ΠΈΡΠ½ΠΈΡ Π±Π΅Π»ΠΊΠΎΠ²ΠΎΠ³ΠΎ ΠΎΠΊΡΡΠΆΠ΅Π½ΠΈΡ Π²Π±Π»ΠΈΠ·ΠΈ ΠΌΠΎΠ»Π΅ΠΊΡΠ» Π±Π°ΠΊΡΠ΅ΡΠΈΠΎΡ Π»ΠΎΡΠΎΡΠΈΠ»Π»ΠΎΠ² Π Π΄ ΠΈ ΠΠ² Π½Π° ΡΠ²ΠΎΠΉΡΡΠ²Π° ΠΈ ΡΡΠ½ΠΊΡΠΈΠΎΠ½Π°Π»ΡΠ½ΡΡ Π°ΠΊΡΠΈΠ²Π½ΠΎΡΡΡ Π Π¦ Rhodobacter sphaeroides. Π ΡΠ²ΡΠ·ΠΈ Ρ ΠΏΠΎΡΡΠ°Π²Π»Π΅Π½Π½ΠΎΠΉ ΡΠ΅Π»ΡΡ Π±ΡΠ»ΠΈ ΠΎΠΏΡΠ΅Π΄Π΅Π»Π΅Π½Ρ ΡΠ»Π΅Π΄ΡΡΡΠΈΠ΅ Π·Π°Π΄Π°ΡΠΈ:
β’ ΠΠ½Π΅ΡΠ΅Π½ΠΈΠ΅ Π½Π°ΠΏΡΠ°Π²Π»Π΅Π½Π½ΡΡ ΠΌΡΡΠ°ΡΠΈΠΉ Π² pufLΠΈ pufMΠ³Π΅Π½Ρ ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΠΎΠ³ΠΎ ΡΠ΅Π½ΡΡΠ° Rba. sphaeroides.
β’ ΠΠΎΠ΄Π±ΠΎΡ ΡΡΠ»ΠΎΠ²ΠΈΠΉ Π΄Π»Ρ Π²ΡΠ΄Π΅Π»Π΅Π½ΠΈΡ Π³Π΅Π½Π΅ΡΠΈΡΠ΅ΡΠΊΠΈ ΠΌΠΎΠ΄ΠΈΡΠΈΡΠΈΡΠΎΠ²Π°Π½Π½ΡΡ Π Π¦ I (L177)H, H (L173)L+ I (L177)H ΠΈ H (M182)L+ I (L177)H ΠΈ ΠΈΡ Π²ΡΠ΄Π΅Π»Π΅Π½ΠΈΠ΅.
β’ ΠΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΠ΅ Π²Π»ΠΈΡΠ½ΠΈΡ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΡΡ Π·Π°ΠΌΠ΅ΡΠ΅Π½ΠΈΠΉ Π² L177, L173 ΠΈ Ml82 ΠΏΠΎΠ·ΠΈΡΠΈΡΡ Π½Π° ΡΠ²ΠΎΠΉΡΡΠ²Π° Π³Π΅Π½Π΅ΡΠΈΡΠ΅ΡΠΊΠΈ. ΠΌΠΎΠ΄ΠΈΡΠΈΡΠΈΡΠΎΠ²Π°Π½Π½ΡΡ Π Π¦ I (L177)H, H (L173)L+ I (L177)H ΠΈ H (M182)L+ I (L177)H.
ΠΠ°ΡΡΠ½Π°Ρ Π½ΠΎΠ²ΠΈΠ·Π½Π°.
Π‘ ΠΏΠΎΠΌΠΎΡΡΡ ΠΌΠ΅ΡΠΎΠ΄Π° ΡΠ°ΠΉΡ-Π½Π°ΠΏΡΠ°Π²Π»Π΅Π½Π½ΠΎΠ³ΠΎ ΠΌΡΡΠ°Π³Π΅Π½Π΅Π·Π° Π±ΡΠ»ΠΈ ΡΠΎΠ·Π΄Π°Π½Ρ Π½ΠΎΠ²ΡΠ΅ ΠΊΠΎΠΌΠ±ΠΈΠ½Π°ΡΠΈΠΈ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΡΡ Π·Π°ΠΌΠ΅ΡΠ΅Π½ΠΈΠΉ I (L177)H, H (L 173) L+I (L 177) Π ΠΈ H (M182)L+ I (L177)H Π² ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΠΎΠΌ ΡΠ΅Π½ΡΡΠ΅ Rba. sphaeroides. ΠΠΎΠ»ΡΡΠ΅Π½Ρ Π Π¦ I (L177)H, Π² ΠΊΠΎΡΠΎΡΡΡ Π°ΡΠΎΠΌ ΠΌΠ°Π³Π½ΠΈΡ ΠΠ₯Π» ΠΠ² 6&trade—ΠΊΠΎΠΎΡΠ΄ΠΈΠ½ΠΈΡΠΎΠ²Π°Π½. ΠΠΎΠΊΠ°Π·Π°Π½ΠΎ, ΡΡΠΎ Π² ΡΠ΅Π·ΡΠ»ΡΡΠ°ΡΠ΅ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΠΎΠ³ΠΎ Π·Π°ΠΌΠ΅ΡΠ΅Π½ΠΈΡ ΠΠ΅.
His Π² Π Π¦ I (L177)H ΠΈ I (L177)H+H (M182)L ΠΏΡΠΎΠΈΠ·ΠΎΡΠ»ΠΎ ΠΊΠΎΠ²Π°Π»Π΅Π½ΡΠ½ΠΎΠ΅ ΡΠ²ΡΠ·ΡΠ²Π°Π½ΠΈΠ΅ i ΠΌΠΎΠ»Π΅ΠΊΡΠ»Ρ ΠΠ₯Π» ΠΈ L-ΡΡΠ±ΡΠ΅Π΄ΠΈΠ½ΠΈΡΡ. ΠΠΎΠ»ΡΡΠ΅Π½Ρ Π Π¦ Π΄Π²ΠΎΠΉΠ½ΠΎΠ³ΠΎ ΠΌΡΡΠ°Π½ΡΠ° I (L177)H+H (M182)L, Π³Π΄Π΅ Π»ΠΈΠ³Π°Π½Π΄ Π°ΡΠΎΠΌΠ° ΠΌΠ°Π³Π½ΠΈΡ ΠΌΠΎΠ»Π΅ΠΊΡΠ»Ρ ΠΠ₯Π» ΠΠ² ΡΠ°ΡΠΏΠΎΠ»ΠΎΠΆΠ΅Π½ Π½Π΅ Ρ ΠΎΠ±ΡΡΠ½ΠΎΠΉ, Π°ΡΡΠΎΡΠΎΠ½Ρ, Π° Ρ (3-ΡΡΠΎΡΠΎΠ½Ρ ΠΌΠ°ΠΊΡΠΎΡΠΈΠΊΠ»Π° ΡΠ΅ΡΡΠ°ΠΏΠΈΡΡΠΎΠ»ΡΠ½ΠΎΠ³ΠΎ ΠΊΠΎΠ»ΡΡΠ°. Π£ΡΡΠ°Π½ΠΎΠ²Π»Π΅Π½ΠΎ, ΡΡΠΎ Π² Π Π¦ H (L 173) L+I (L 177) Π, Π² ΠΎΡΡΡΡΡΡΠ²ΠΈΠ΅ ΠΏΡΠΈΡΠΎΠ΄Π½ΠΎΠ³ΠΎ Π»ΠΈΠ³Π°Π½Π΄Π° Π°ΡΠΎΠΌΠ° ΠΌΠ°Π³Π½ΠΈΡ — Π³ΠΈΡΡΠΈΠ΄ΠΈΠ½Π° L173, Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΡΠΉ ΠΎΡΡΠ°ΡΠΎΠΊ Π³ΠΈΡΡΠΈΠ΄ΠΈΠ½Π° L177 Π»ΠΈΠ³Π°Π½Π΄ΠΈΡΡΠ΅Ρ Π°ΡΠΎΠΌ ΠΌΠ°Π³Π½ΠΈΡ Π±Π°ΠΊΡΠ΅ΡΠΈΠΎΡ Π»ΠΎΡΠΎΡΠΈΠ»Π»Π° Π Π.
ΠΠΏΡΠΎΠ±Π°ΡΠΈΡ ΡΠ°Π±ΠΎΡΡ.
ΠΠ°ΡΠ΅ΡΠΈΠ°Π»Ρ Π΄ΠΈΡΡΠ΅ΡΡΠ°ΡΠΈΠΈ Π±ΡΠ»ΠΈ ΠΏΡΠ΅Π΄ΡΡΠ°Π²Π»Π΅Π½Ρ Π½Π° 9-ΠΎΠΉ ΠΈ 11 ΠΌΠ΅ΠΆΠ΄ΡΠ½Π°ΡΠΎΠ΄Π½ΡΡ ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΡΡ ΠΌΠΎΠ»ΠΎΠ΄ΡΡ ΡΡΠ΅Π½ΡΡ (ΠΡΡΠΈΠ½ΠΎ, 2005; 2007), ΠΠ΅ΠΆΠ΄ΡΠ½Π°ΡΠΎΠ΄Π½ΠΎΠΌ Π±ΠΈΠΎΡΠΈΠ·ΠΈΡΠ΅ΡΠΊΠΎΠΌ ΠΊΠΎΠ½Π³ΡΠ΅ΡΡΠ΅ (Π€ΡΠ°Π½ΡΠΈΡ, 2005), XVIII ΠΡΡΠΈΠ½ΡΠΊΠΈΡ ΡΡΠ΅Π½ΠΈΡΡ ΠΏΠΎ ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅Π·Ρ ΠΈ ΠΡΠ΅ΡΠΎΡΡΠΈΠΉΡΠΊΠΎΠΉ ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΠΈ «ΠΡΠ΅ΠΎΠ±ΡΠ°Π·ΠΎΠ²Π°Π½ΠΈΠ΅ ΡΠ½Π΅ΡΠ³ΠΈΠΈ ΡΠ²Π΅ΡΠ° ΠΏΡΠΈ ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅Π·Π΅ (ΠΡΡΠΈΠ½ΠΎ, 2005), ΠΠ° ΠΌΠ΅ΠΆΠ΄ΡΠ½Π°ΡΠΎΠ΄Π½ΠΎΠΉ ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΠΈ «Π€ΠΎΡΠΎΡΠΈΠ½ΡΠ΅Π· Π² ΠΏΠΎΡΡ-Π³Π΅Π½ΠΎΠΌΠ½ΡΡ ΡΡΡ» (ΠΡΡΠΈΠ½ΠΎ,.
2006), ΠΠ΅ΠΆΠ΄ΡΠ½Π°ΡΠΎΠ΄Π½ΠΎΠΌ ΠΊΠΎΠ½Π³ΡΠ΅ΡΡΠ΅ ΠΏΠΎ Π±ΠΈΠΎΡ ΠΈΠΌΠΈΠΈ ΠΈ ΠΌΠΎΠ»Π΅ΠΊΡΠ»ΡΡΠ½ΠΎΠΉ Π±ΠΈΠΎΠ»ΠΎΠ³ΠΈΠΈ (Π―ΠΏΠΎΠ½ΠΈΡ, 2006), ΠΠ΅ΠΆΠ΄ΡΠ½Π°ΡΠΎΠ΄Π½ΠΎΠΌ ΡΠΈΠΌΠΏΠΎΠ·ΠΈΡΠΌΠ΅ ΠΏΠΎ ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΈΠΌ ΠΏΡΠΎΠΊΠ°ΡΠΈΠΎΡΠ°ΠΌ (Π€ΡΠ°Π½ΡΠΈΡ, 2006), ΠΠ΅ΠΆΠ΄ΡΠ½Π°ΡΠΎΠ΄Π½ΠΎΠΉ ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΠΈ ΠΏΠΎ ΡΠ΅ΡΡΠ°ΠΏΠΈΡΡΠΎΠ»ΡΠ½ΡΠΌ ΡΠΎΡΠΎΡΠ΅ΡΠ΅ΠΏΡΠΎΡΠ°ΠΌ ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅ΡΠΈΡΠ΅ΡΠΊΠΈΡ ΠΎΡΠ³Π°Π½ΠΈΠ·ΠΌΠΎΠ² (Π―ΠΏΠΎΠ½ΠΈΡ,.
2007), ΠΠ²ΡΠΎΠΏΠ΅ΠΉΡΠΊΠΎΠΉ Π±ΠΈΠΎΡΠ½Π΅ΡΠ³Π΅ΡΠΈΡΠ΅ΡΠΊΠΎΠΉ ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΠΈ (ΠΡΠ»Π°Π½Π΄ΠΈΡ, 2008), ΠΠ΅ΠΆΠ΄ΡΠ½Π°ΡΠΎΠ΄Π½ΠΎΠΉ ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΠΈ «ΠΡΠ΅ΠΎΠ±ΡΠ°Π·ΠΎΠ²Π°Π½ΠΈΠ΅ ΡΠ½Π΅ΡΠ³ΠΈΠΈ ΡΠ²Π΅ΡΠ° ΠΏΡΠΈ ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅Π·Π΅» (ΠΡΡΠΈΠ½ΠΎ, 2008), Π½Π° ΠΠΎΡΠ΄ΠΎΠ½ΠΎΠ²ΡΠΊΠΈΡ ΡΡΠ΅Π½ΠΈΡΡ ΠΏΠΎ ΡΠΎΡΠΎΡΠΈΠ½ΡΠ΅Π·Ρ.
Π‘Π¨Π, 2008), Π½Π° ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΠΈ «Π€ΠΎΡΠΎΡ ΠΈΠΌΠΈΡ Ρ Π»ΠΎΡΠΎΡΠΈΠ»Π»Π° Π² ΠΌΠΎΠ΄Π΅Π»ΡΠ½ΡΡ ΠΈ ΠΏΡΠΈΡΠΎΠ΄Π½ΡΡ ΡΠΈΡΡΠ΅ΠΌΠ°Ρ » (ΠΡΡΠΈΠ½ΠΎ, 2009), ΠΈ Π½Π° XIII ΠΠ²ΡΠΎΠΏΠ΅ΠΉΡΠΊΠΎΠΉ ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΠΈ ΠΏΠΎ ΡΠΏΠ΅ΠΊΡΡΠΎΡΠΊΠΎΠΏΠΈΠΈ Π±ΠΈΠΎΠ»ΠΎΠ³ΠΈΡΠ΅ΡΠΊΠΈΡ ΠΌΠΎΠ»Π΅ΠΊΡΠ» (ΠΡΠ°Π»ΠΈΡ, 2009).
Π‘Π²ΡΠ·Ρ Ρ ΠΏΠ»Π°Π½ΠΎΠΌ Π½Π°ΡΡΠ½ΠΎ-ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°ΡΠ΅Π»ΡΡΠΊΠΎΠ³ΠΎ ΠΈΠ½ΡΡΠΈΡΡΡΠ°.
Π’Π΅ΠΌΠ° Π΄ΠΈΡΡΠ΅ΡΡΠ°ΡΠΈΠΈ ΡΠ²Π»ΡΠ΅ΡΡΡ ΡΠ°ΡΡΡΡ ΠΏΠ»Π°Π½ΠΎΠ²ΠΎΠΉ ΡΠ΅ΠΌΡ ΠΈΠ½ΡΡΠΈΡΡΡΠ°. Π Π°Π±ΠΎΡΠ° Π²ΡΠΏΠΎΠ»Π½ΡΠ»Π°ΡΡ ΠΏΡΠΈ ΡΠΈΠ½Π°Π½ΡΠΎΠ²ΠΎΠΉ ΠΏΠΎΠ΄Π΄Π΅ΡΠΆΠΊΠ΅ Π€Π΅Π΄Π΅ΡΠ°Π»ΡΠ½ΡΠΌ Π°Π³Π΅Π½ΡΡΡΠ²ΠΎΠΌ ΠΏΠΎ Π½Π°ΡΠΊΠ΅ ΠΈ ΠΈΠ½Π½ΠΎΠ²Π°ΡΠΈΡΠΌ Π ΠΎΡΡΠΈΠΉΡΠΊΠΎΠΉ Π°ΠΊΠ°Π΄Π΅ΠΌΠΈΠΈ Π½Π°ΡΠΊ (02.512.11.2085; 02.740.11.0293), ΠΏΡΠΎΠ³ΡΠ°ΠΌΠΌΠ°ΠΌΠΈ ΠΡΠ΅Π·ΠΈΠ΄ΠΈΡΠΌΠ° Π ΠΠ «ΠΠΎΠ»Π΅ΠΊΡΠ»ΡΡΠ½Π°Ρ ΠΈ ΠΊΠ»Π΅ΡΠΎΡΠ½Π°Ρ Π±ΠΈΠΎΠ»ΠΎΠ³ΠΈΡ», «ΠΠΊΡΡΡΠ΅ΠΌΠ°Π»ΡΠ½ΡΠ΅ ΡΠ²Π΅ΡΠΎΠ²ΡΠ΅ ΠΏΠΎΠ»Ρ ΠΈ ΠΈΡ ΠΏΡΠΈΠ»ΠΎΠΆΠ΅Π½ΠΈΠ΅», Π³ΡΠ°Π½ΡΠΎΠΌ ΠΡΠ΅Π·ΠΈΠ΄Π΅Π½ΡΠ° Π Π€ ΠΠ¨-3277.2006.4- ΠΠ¨-4525.2008.4) ΠΈ Π Π€Π€Π (03−04−48 215- 06−04−48 686- 09−04−109).
ΠΡΠ±Π»ΠΈΠΊΠ°ΡΠΈΠΈ.
ΠΠΎ ΠΌΠ°ΡΠ΅ΡΠΈΠ°Π»Π°ΠΌ Π΄ΠΈΡΡΠ΅ΡΡΠ°ΡΠΈΠΈ ΠΎΠΏΡΠ±Π»ΠΈΠΊΠΎΠ²Π°Π½ΠΎ 27 ΡΠ°Π±ΠΎΡ, ΠΈΠ· Π½ΠΈΡ ΠΎΠ΄Π½Π° ΡΡΠ°ΡΡΡ Π² Π·Π°ΡΡΠ±Π΅ΠΆΠ½ΠΎΠΌ ΠΆΡΡΠ½Π°Π»Π΅ FEBS letters, ΡΡΠΈ ΡΡΠ°ΡΡΠΈ Π² ΠΎΡΠ΅ΡΠ΅ΡΡΠ²Π΅Π½Π½ΠΎΠΌ ΠΆΡΡΠ½Π°Π»Π΅ ΠΠΈΠΎΡ ΠΈΠΌΠΈΡ, 23 ΡΠ΅Π·ΠΈΡΠ° Π² ΠΌΠ°ΡΠ΅ΡΠΈΠ°Π»Π°Ρ ΠΎΡΠ΅ΡΠ΅ΡΡΠ²Π΅Π½Π½ΡΡ ΠΈ ΠΌΠ΅ΠΆΠ΄ΡΠ½Π°ΡΠΎΠ΄Π½ΡΡ ΠΊΠΎΠ½ΡΠ΅ΡΠ΅Π½ΡΠΈΠΉ.
Π‘ΡΡΡΠΊΡΡΡΠ° ΠΈ ΠΎΠ±ΡΠ΅ΠΌ ΡΠ°Π±ΠΎΡΡ.
ΠΠΈΡΡΠ΅ΡΡΠ°ΡΠΈΡ ΡΠΎΡΡΠΎΠΈΡ ΠΈΠ· Π²Π²Π΅Π΄Π΅Π½ΠΈΡ, ΠΎΠ±Π·ΠΎΡΠ° Π»ΠΈΡΠ΅ΡΠ°ΡΡΡΡ, ΠΎΠΏΠΈΡΠ°Π½ΠΈΡ ΠΌΠ°ΡΠ΅ΡΠΈΠ°Π»ΠΎΠ² ΠΈ ΠΌΠ΅ΡΠΎΠ΄ΠΎΠ² ΠΈΡΡΠ»Π΅Π΄ΠΎΠ²Π°Π½ΠΈΡ, ΠΈΠ·Π»ΠΎΠΆΠ΅Π½ΠΈΡ ΡΠ΅Π·ΡΠ»ΡΡΠ°ΡΠΎΠ² ΠΈ ΠΈΡ ΠΎΠ±ΡΡΠΆΠ΄Π΅Π½ΠΈΡ, Π·Π°ΠΊΠ»ΡΡΠ΅Π½ΠΈΡ, Π²ΡΠ²ΠΎΠ΄ΠΎΠ² ΠΈ ΡΠΏΠΈΡΠΊΠ° ΡΠΈΡΠΈΡΡΠ΅ΠΌΠΎΠΉ Π»ΠΈΡΠ΅ΡΠ°ΡΡΡΡ. ΠΠΈΡΡΠ΅ΡΡΠ°ΡΠΈΡ ΠΈΠ·Π»ΠΎΠΆΠ΅Π½Π° Π½Π° 153 ΡΡΡΠ°Π½ΠΈΡΠ°Ρ , ΠΈΠ»Π»ΡΡΡΡΠΈΡΠΎΠ²Π°Π½Π° 49 ΡΠΈΡΡΠ½ΠΊΠ°ΠΌΠΈ ΠΈ 5 ΡΠ°Π±Π»ΠΈΡΠ°ΠΌΠΈ.
Π‘ΠΏΠΈΡΠΎΠΊ Π»ΠΈΡΠ΅ΡΠ°ΡΡΡΡ
ΡΠΎΠ΄Π΅ΡΠΆΠΈΡ 190 ΠΈΡΡΠΎΡΠ½ΠΈΠΊΠΎΠ² (ΠΈΠ· Π½ΠΈΡ 178 Π½Π° Π°Π½Π³Π»ΠΈΠΉΡΠΊΠΎΠΌ ΡΠ·ΡΠΊΠ΅).
ΠΠ«ΠΠΠΠ«:
1. Π ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΡΠ΅ ΡΠ΅Π½ΡΡΡ Rba. sphaeroides Π²Π±Π»ΠΈΠ·ΠΈ ΠΌΠΎΠ»Π΅ΠΊΡΠ» ΠΠ₯Π» Π Π ΠΈ ΠΠ² Π²Π²Π΅Π΄Π΅Π½Ρ Π½ΠΎΠ²ΡΠ΅ ΠΊΠΎΠΌΠ±ΠΈΠ½Π°ΡΠΈΠΈ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΡΡ Π·Π°ΠΌΠ΅ΡΠ΅Π½ΠΈΠΉ I (L177)H, H (L 173) L+I (L 177) Π, H (M182)L+I (L177)H.
2. ΠΠΎΠΊΠ°Π·Π°Π½ΠΎ, ΡΡΠΎ Π² ΡΠ΅Π·ΡΠ»ΡΡΠ°ΡΠ΅ ΠΌΡΡΠ°ΡΠΈΠΈ I (L177)H Π² Π Π¦ Rba. sphaeroides Π°ΡΠΎΠΌ ΠΌΠ°Π³Π½ΠΈΡ ΠΠ₯Π» ΠΠ² ΡΡΠ°Π½ΠΎΠ²ΠΈΡΡΡ ΡΠ΅ΡΡΠΈ ΠΊΠΎΠΎΡΠ΄ΠΈΠ½ΠΈΡΠΎΠ²Π°Π½Π½ΡΠΌ, ΡΡΠΎ ΠΏΡΠΈΠ²ΠΎΠ΄ΠΈΡ ΠΊ ΠΏΠΎΡΠ²Π»Π΅Π½ΠΈΡ ΠΏΠΎΠ»ΠΎΡΡ 638 Π½ΠΌ Π² Π²ΠΈΠ΄ΠΈΠΌΠΎΠΉ ΠΎΠ±Π»Π°ΡΡΠΈ ΡΠΏΠ΅ΠΊΡΡΠ° ΠΏΠΎΠ³Π»ΠΎΡΠ΅Π½ΠΈΡ Π Π¦.
3. ΠΠΏΠ΅ΡΠ²ΡΠ΅ ΠΏΠΎΠΊΠ°Π·Π°Π½ΠΎ, ΡΡΠΎ Π² ΡΠ΅Π·ΡΠ»ΡΡΠ°ΡΠ΅ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΠΎΠ³ΠΎ Π·Π°ΠΌΠ΅ΡΠ΅Π½ΠΈΡ I (L177)H ΠΎΠ±ΡΠ°Π·ΡΠ΅ΡΡΡ ΠΊΠΎΠ²Π°Π»Π΅Π½ΡΠ½Π°Ρ ΡΠ²ΡΠ·Ρ ΠΌΠ΅ΠΆΠ΄Ρ ΠΌΠΎΠ»Π΅ΠΊΡΠ»ΠΎΠΉ ΠΠ₯Π» Π Π ΠΈ L-ΡΡΠ±ΡΠ΅Π΄ΠΈΠ½ΠΈΡΠ΅ΠΉ Π Π¦.
4. Π£ΡΡΠ°Π½ΠΎΠ²Π»Π΅Π½ΠΎ, ΡΡΠΎ Π² Π³Π΅Π½Π΅ΡΠΈΡΠ΅ΡΠΊΠΈ ΠΌΠΎΠ΄ΠΈΡΠΈΡΠΈΡΠΎΠ²Π°Π½Π½ΡΡ Π Π¦ H (M182)L+I (L177)H Π½Π°Π±Π»ΡΠ΄Π°Π΅ΡΡΡ ΡΠ»ΡΡΠ°ΠΉ |3-Π»ΠΈΠ³Π°Π½Π΄ΠΈΡΠΎΠ²Π°Π½ΠΈΡ Π°ΡΠΎΠΌΠ° ΠΌΠ°Π³Π½ΠΈΡ Π±Π°ΠΊΡΠ΅ΡΠΈΠΎΡ Π»ΠΎΡΠΎΡΠΈΠ»Π»Π° Π² ΡΠ΅Π°ΠΊΡΠΈΠΎΠ½Π½ΡΡ ΡΠ΅Π½ΡΡΠ°Ρ ΠΏΡΡΠΏΡΡΠ½ΡΡ Π±Π°ΠΊΡΠ΅ΡΠΈΠΉ.
5. ΠΠΎΠΊΠ°Π·Π°Π½ΠΎ, ΡΡΠΎ Π² Π Π¦ H (L173)L+I (L177)H ΡΠΌΠ΅ΡΠ΅Π½ΠΈΠ΅ Π°ΠΌΠΈΠ½ΠΎΠΊΠΈΡΠ»ΠΎΡΠ½ΠΎΠ³ΠΎ ΠΎΡΡΠ°ΡΠΊΠ° Π³ΠΈΡΡΠΈΠ΄ΠΈΠ½Π° — Π»ΠΈΠ³Π°Π½Π΄Π° Π°ΡΠΎΠΌΠ° ΠΌΠ°Π³Π½ΠΈΡ ΠΠ₯Π» Π Π Ρ L173 Π½Π° L177 ΠΏΠΎΠ·ΠΈΡΠΈΡ ΠΏΡΠΈΠ²ΠΎΠ΄ΠΈΡ ΠΊ 45-Π½ΠΌ ΠΊΠΎΡΠΎΡΠΊΠΎΠ²ΠΎΠ»Π½ΠΎΠ²ΠΎΠΌΡ ΡΠ΄Π²ΠΈΠ³Ρ ΠΌΠ°ΠΊΡΠΈΠΌΡΠΌΠ° ΠΏΠΎΠ»ΠΎΡΡ Π , Π½ΠΎ Π½Π΅ ΠΎΠΊΠ°Π·ΡΠ²Π°Π΅Ρ Π·Π°ΠΌΠ΅ΡΠ½ΠΎΠ³ΠΎ Π²Π»ΠΈΡΠ½ΠΈΡ Π½Π° ΡΠ»Π΅ΠΊΡΡΠΎΡ ΠΈΠΌΠΈΡΠ΅ΡΠΊΠΈΠ΅ ΡΠ²ΠΎΠΉΡΡΠ²Π° ΠΏΠ΅ΡΠ²ΠΈΡΠ½ΠΎΠ³ΠΎ Π΄ΠΎΠ½ΠΎΡΠ° ΡΠ»Π΅ΠΊΡΡΠΎΠ½Π° ΠΈ ΡΠΎΡΠΎΡ ΠΈΠΌΠΈΡΠ΅ΡΠΊΡΡ Π°ΠΊΡΠΈΠ²Π½ΠΎΡΡΡ.
Π Π¦.
Π‘ΠΏΠΈΡΠΎΠΊ Π»ΠΈΡΠ΅ΡΠ°ΡΡΡΡ
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